Description
Key Characters:
Growth Form:
Shrubs, creeping and sometimes rooting at the nodes or erect, sometimes trees, often foetid when bruised.
Stems:
Roots:
Leaves:
Leaves simple.
Opposite or rarely ternate.
Margins entire.
Petiolate or sessile.
Stipules interpetiolar, distinct or partly connate, entire or dentate with teethlike marginal appendages (in Hawai‘i these reddish brown to reddish purple; usually bearing mucilage-producing colleters on inner surface that aid in protection of the growing shoot tip.
Flowers:
Flowers axillary, solitary or in cymes.
Flowers insect-pollinated, unisexual (and the plants dioecious), rarely polygamous or 1 species bisexual (perfect), often heterostylous.
Calyx 4–5(–10)-toothed with open aestivation, usually absent in staminate flowers.
Corolla actinomorphic, rarely irregular or zygomorphic and bilabiate, (3)4-5(8-10)-lobed, the lobes valvate, imbricate, or convolute; nectary disk usually present.
Stamens 4-5(-10), alternate with the corolla lobes, inserted at base of corolla tube; filaments long-exserted; anthers penduous, dithecal, opening by longitudinal slits.
Ovary inferior, occasionally partly so, or very rarely superior, 2(-4)-celled; ovule 1 per cell, basal, anatropous; style as many as carpels, 2(-4)lobed, divided nearly to base, terminal, slender; stigmas lobed or capitate, long-exserted, papillose-hirsute, dry or occasionally wet.
Fruit:
Fruit drupaceous; juicy; ovoid to globose; with 2(-4); 1-seeded; planoconvex pyrenes.
Seeds with well-developed oily endosperm; or endosperm occasionally scanty or absent.
Ploidy:
Habitat:
Elevation Range: